Research Interests:
Forest Sciences Centre
2424 Main Mall
Vancouver, BC V6T 1Z4
work phone: 6048228101

Research Areas

population genetics
plant mating systems


Unit Associations

Beaty Biodiversity Research Centre
Centre for Forest Gene Conservation CFGC
Department of Botany
Genetic Data Center GDC

Professional Affiliations

AOB Plants, Associate Editor
Genetics Society of Britain – Heredity, Associate Editor

Current Graduate Students

Gwendolyn Huber, PhD
Chemical ecology of blueberries in relation to bee pollination
Simon Nadeau, MSc
Adaptive genetic variation in Eastern and Western White pine

Post Doctoral Fellows and Research Assistants

Hailin Liu

Current Courses

Winter 2016

FRST432 Molecular Ecology Sections

Use of molecular genetic tools to investigate ecological processes in natural population; DNA variation and its application in landscape genetics, phylogeography, behavioural ecology, conservation genetics, and association genetics. Emphasis placed on statistical and computational data analysis.

Winter 2016

FRST510 Applied Population Genetics Sections

Winter 2016

CONS449C Directed Studies in Natural Resources Conservation - DIR ST NTL RS CN Sections

In special cases and with the approval of the instructor concerned, a student may carry out directed studies of specific problems in natural resources conservation.

Other Courses and Workshops

CONS302 – Issues in Genomics and the Environment, new course started Fall 2015

Selected Publications

Huang, A., J. E. Elliott, K. M. Cheng, K. Ritland, C. Ritland, S. K. Thompson, S. Hindmarch, and K. Martin (2016). Barn owls (Tyto alba) in North America: Phylogeographic structure, connectivity, and genetic diversity. Conservation Genetics 17: 357-367
Huang AC, Elliott JE, Cheng KMT, Ritland K, Ritland CE, Thomsen SK, Hindmarch S, Martin K (2016). Barn owls (Tyto alba) in North America: Phylogeographic structure, connectivity, and genetic diversity Conservation Genetics 17: 357-367
Baoguo L, Payseur B; Huang K, Ritland K; Dunn DW, Qi Z, Guo G (2016). Estimating relatedness with the presence of null alleles Genetics 202: 247-260
Napolitano C, Díaz D, Johnson WE, Sanderson J, Ritland K, Ritland CE, Poulin E (2016). Reduced genetic diversity and increased dispersal in Guigna (Leopardus guigna) in Chilean fragmented landscapes Journal of Heredity 106: 522–536
Nadeau S, Godbout J, Lamothe M, Gros-Louis M-C, Isabel N, Ritland K. (2015). Contrasting patterns of genetic diversity across the ranges of Pinus monticola and P. strobus: A comparison between eastern and western North American postglacial colonization histories. American Journal of Botany 102: 1342-1355
Huang K, Ritland K, Guo S, Chen D, Chen S, Ren Y, He G, Li B (2015). Estimating pairwise relatedness between individuals with different levels of ploidy. Molecular Ecology Resources 15: 772-784
De La Torre A, Birol I, Bousquet J, Ingvarsson PK, Jansson S, Jones S, Keeling C, MacKay J, Nilsson O, Ritland K, Street N, Yanchuk A, Zerbe P, Bohlmann J (2015). Insights into conifer giga-genomes Plant Physiology 166: 1724-1732
Napolitano C, Johnson WE, Sanderson J, O’Brien SJ, Hoelzel AR, Freer R, Dunstone N, Ritland K, Ritland CE, Poulin E. (2014). Phylogeography and population history of Leopardus guigna, the smallest American felid Conservation Genetics 15: 631-653
Sena JS, Sena JS, Giguere I, Boyle B, Rigault P, Birol I, Zuccolo A, Ritland K, Ritland C, Bohlmann J, Jones S, Bousquet J, Mackay J (2014). Evolution of gene structure in the conifer Picea glauca: A comparative analysis of the impact of intron size BMC Plant Biology 14: 95 (16 April 2014)
Huang K, Ritland K, Guo S, Shattuck M, Li B. (2014). A pair-wise relatedness estimator for tetraploids Molecular Ecology Resources 14: 734-744
Chen C, Ritland K (2013). Lineage-specific mapping of quantitative trait loci Heredity 111: 106-113
Birol I, Raymond A, Jackman SD, Pleasance S, Coope R, Taylor GA, Saint Yuen MM, Keeling CI, Brand D, Vandervalk BP, Kirk H, Pandoh P, Moore RA, Zhao Y, Mungall AJ, Jaquish B, Yanchuk A, Ritland C, Boyle B, Bousquet j, Ritland K, MacKay J, Bohlmann J, Jones SJM (2013). Assembling the 20 Gb white spruce (Picea glauca) genome from whole-genome shotgun sequencing data Bioinformatics 29: 1492-1497
Nystedt B, Street NR, Wetterbom A, Zuccolo A, Lin Y-C, Scofield DG, Vezzi F, Delhomme N, Giacomello S, Alexeyenko A, Vicedomini R, Sahlin K, Sherwood E, Elfstand M, Gramzow L, Holmberg K, Hällman J, Keech O, Klasson L, Koriabine M, Kucukoglu M, Käller M, Luthman J, Lysholm F, Niittylä T, Olson A, Rilakovic N, Ritland C, Rosselló JA, Sena J, Svensson T, Talavera-López C, Theißen G, Tuominen H, Vanneste K, Wu Z-Q, Zhang B, Zerbe P, Arvestad L, Bhalerao R, Bohlmann J, Bousquet J, Garcia-Gil R, Hvidsten TR, de Jong P, MacKay J, Morgante M, Ritland K, Sundberg B, Thompson SL, Van de Peer Y, Andersson B, Nilsson O, Ingvarsson PK, Lundeberg J, Jansson S (2013). The Norway spruce genome sequence gives insights into conifer genome evolution Nature (in press)
Porth I, White R, Jaquish B, Alfaro R, Ritland C, Ritland K (2013). Genetical genomics identifies the genetic architecture for growth and weevil resistance in spruce PloS one 7 (9), e44397
Napolitano C, Sanderson J, Johnson W, O’Brien S, Hoelzel A, Freer R, Dunstone N, Ritland K, Poulin E (2012). Population genetics of the felid Leopardus guigna in Southern South America: Identifying inbraspecific units for conservation Nova Science Publisher, Inc. Hauppauge NY USA Chapter 11, In: Molecular Population Genetics, Phylogenetics, Evolutionary Biology and Conservation of the Neotropical Carnivores Editors: Manuel Ruiz-García and Joseph Shostell
Ritland, Kermit (2012). Genomics of a phylum distant from flowering plants: conifers. Tree Genetics and Genomics 8:573-582
Colin T. Kelleher C, Wilkin J, Zhuang J, Javier Cortés J, Quintero A, Gallagher T, Bohlmann J, Douglas C, Ellis B, Ritland K. (2012). SNP discovery, gene diversity, and linkage disequilibrium in wild populations of Populus tremuloides. Tree Genetics and Genomics 8: 821-829.
Emmanuel Buschiazzo Carol Ritland Jorg Bohlmann Kermit Ritland (2012). Slow but not low: genomic comparisons reveals slower evolutionary rate and higher dN/dS in conifers compared to angiosperms BMC Evolutionary Biology 2012, 12:8 doi:10.1186/1471-2148-12-8
Hedrick P, Ritland K. (2012). Population genetics of the white-phased `spirit`black bear of British Columbia. Evolution 66: 305–313, February 2012
Kremer A, Ronce O, Robledo-Arnuncio J, Guillaume F, Bohrer G, Nathan R, Bridle J, Gomulkiewicz R, Klein E, Ritland K, Kuparinen A, Gerber S, Schueler S (2012). Long-distance gene flow and adaptation of forest trees to rapid climate change Ecology Letters 15: 378-392
Verne S, Jaquish B, White R, Ritland C, Ritland K (2011). Global transcriptome analysis of constitutive resistance to the White Pine weevil in spruce Genome Biol Evol 3: 851-867
Porth I, Hamberger B, White R, Ritland K (2011). Defense mechanisms against herbivory in Picea: sequence evolution and expression regulation of gene family members in the phenylpropanoid pathway BMC Genomics 12: 608 (26 pages)
Ritland K (2011). Evolutionary potential in the wild: more than meets the eye Molecular Ecology 20: 3494-3495
Pelgas B, Bousquet J, Meirmans PG, Ritland K, Isabel N (2011). QTL mapping in white spruce: gene maps and genomic regions underlying adaptive traits across pedigrees, years and environments BMC Genomics 12: 145 (23 pages)
Ritland K, Krutovsky K, Tsumura Y, Pelgas B, Isabel N, Bousquet J (2011). Genetic mapping in conifers Science Publishers, Enfield, NH USA Chapter 5, pps. 196-238. In Genetics, Genomics and Breeding of Conifers, Plomion C, Bousquet J, Kole C (Eds.)
Holliday, J.A., M. Yuen, K.Ritland and S.N. Aitken (2010). Postglacial history of a widespread conifer produces inverse clines in selective neutrality tests Molecular Ecology 18: 3857-3864.
Holliday, J.A., K. Ritland and S.N. Aitken (2010). Widespread ecologically-relevant genetic markers developed from association mapping of climate-related traits in Sitka spruce (Picea sitchensis) New Phytologist 188: 501-514.
Ally D. Ritland K, Otto SP (2010). Aging in a Long-Lived Clonal Tree PLOS Biology 8: e1000454
Albouyeh R, Farzaneh N, Bohlmann J, Ritland K (2010). Multivariate analysis of digital gene expression profiles identifies a xylem signature of the vascular tissue of white spruce (Picea glauca) Tree Genetics and Genomics. 6: 601-611
Lo EYY, Stefanovic S, Ritland K, Dickinson TA (2010). Fine-scale comparisons of genetic variability in seed families of asexually and sexually reproducing Crataegus (Hawthorn; Rosaceae) American Journal of Botany 97: 1014-1024
Ramsay H, Rieseberg L, Ritland K (2009). Correlation of evolutionary rate with pathway position in plant terpenoid biosynthesis. Molecular Biology and Evolution 26 : 1045-1053
Liewlaksaneeyanawin C, Zhuang J, Tang M, Farzaneh N, Lueng G, Cullis C, Findlay S, Ritland C, Bohlmann J, Ritland K. (2009). Identification of COS markers in the Pinaceae Tree Genetics and Genomes 5: 247-255
Lippert DN, Ralph GS, Philips M, White R, Smith D, Hardie D, Gershenzon J, Ritland K, Borchers CH Bohlmann J. (2009). Quantitative iTRAQ proteome and comparative transcriptome: Analysis of elicitor-Induced Norway Spruce (Picea abies) cells reveals elements of calcium signaling in the early conifer defense response Proteomics 9:350-67
Albouyeh R, Ritland K (2009). Estimating heritability of gene expression using parent-offspring regression with two-channel microarrays J. Heredity 100:114-118.
Ally D, Ritland K, Otto S (2009). Can clone size serve as a proxy for clone age? An exploration using microsatellite divergence in Populus tremuloides Molecular Ecology17: 4897-4911
Björn Hamberger Dawn Hall Mack Yuen Claire Oddy Britta Hamberger Christopher Keeling Carol Ritland Kermit Ritland Jörg Bohlmann (2009). Targeted isolation, sequence assembly and characterization of two white spruce (Picea glauca) BAC clones for terpenoid synthase and cytochrome P450 genes involved in conifer defence reveal insights into a conifer genome BMC Plant Biology 9:106.
Ralph SF, Chun HJE, Kolosova N, Cooper D, Oddy C, Ritland C, Kirkpatrick R, Moore R, Barber S, Holt RA, Jones SJM, Marra MA, Douglas CJ, Ritland K, Bohlmann J. (2008). A conifer genomics resource of 200,000 spruce (Picea spp.) ESTs and 6,464 high-quality, sequence-finished full-length cDNAs for Sitka spruce (Picea sitchensis) BMC Genomics 2008, 9:484
Thompson SL, Bérubé Y, Bruneau A, Ritland K (2008). Three-gene identity coefficients demonstrate that clonal reproduction promotes inbreeding and spatial relatedness in yellow-cedar, Callitropsis nootkatensis. Evolution 62: 2570-2579
O’Connell L , Ritland K, Thompson SL (2008). Post-glacial colonization of western redcedar (Thuja plicata, Cupressaceae) revealed by microsatellite markers Botany 86: 194-203
Stacey Lee Thompson, Gina Choe, Kermit Ritland, and Jeannette Whitton (2008). Cryptic sex within male-sterile polyploid populations of the Easter daisy, Townsendia hookeri International Journal of Plant Sciences169:183–193
Ukrainetz, N. K., Ritland, K. Mansfield, S. D. (2008). Identification of quantitative trait loci for wood quality and growth across eight full-sib coastal Douglas-fir families Tree Genetics and Genomics. 4:159-170
Ukrainetz, N. K., Ritland, K. Mansfield, S. D. (2008). An AFLP linkage map for Douglas-fir based upon multiple full-sib families. Tree Genetics and Genomics. 4:181-191
Kelleher, Colin, Heesun Shin, Ian Bosdet, Chris Fjell, Rradman Chiu, Martin Krzywinski, Jennifer Wilkin, TongMing Yin, Stephen DiFazio, Jennifer Asano, Jun Zhuang, Nicholas Putnam, Steven Jones, Marco Marra, Gerald Tuskan, Joerg Bohlmann, Brian Ellis, Kermit Ritland, Carl Douglas, Jacqueline Schein (2007). A physical map of the highly heterozygous Populus genome: integration with the genome sequence and genetic map The Plant Journal
Lippert D, Chowrira S, Ralph SG, Zhuang J, Aeschliman D, Ritland C, Ritland K, Bohlmann J. (2007). Conifer defense against insects: Proteome analysis of Sitka spruce (Picea sitchensis) bark induced by mechanical wounding or feeding by white pine weevils (Pissodes strobi) Proteomics 7: 248-270
Friedmann M, Ralph SG, Aeschliman D, Zhuang J, Ritland K, Ellis BE, Bohlmann J, and CJ Douglas (2007). Microarray gene expression profiling of the developmental transition from primary to secondary growth and constitutive defence in Sitka spruce (Picea sitchensis) apical shoots Journal of Experimental Botany 58: 593-614
Bousquet J, Isabel N, Pelgas B, Cottrell J, Rungis D, Ritland K (2007). Spruce Springer-Verlag Berlin Heidelberg In Genome Mapping and Molecular Breeding in Plants, Volume 7 Forest Trees C. Kole (Ed.)
Kim S.H., Cheng K.M-T., Ritland C, Ritland K, Silversides FG (2007). Inbreeding in Japanese Quail estimated by pedigree and microsatellite analyses J. Heredity 98: 378-381
Yanik Bérubé, Jun Zhuang, Dainis Rungis, Steven Ralph, Jörg Bohlmann, and Kermit Ritland (2007). Characterization of EST-SSRs in loblolly pine and spruce Tree genetics and genomes 3: 251-259
Ralph S, Oddy C, Cooper D, Yueh H, Jancsik S, Kolosova N, Philippe RN, Aeschliman D, White R, Huber D, Ritland CE, Benoit F, Rigby T, Nantel A, Butterfield YSN, Kirkpatrick R, Chun E, Liu J, Palmquist D, Wynhoven B, Stott J, Yang G, Barber S, Holt RA, Siddiqui A, Jones SJM, Marra MA, Ellis BE, Douglas CJ, Ritland K and Bohlmann J (2006). Genomics of hybrid poplar (Populus trichocarpa x deltoides) interacting with forest tent caterpillars (Malacosoma disstria): Normalized and full-length cDNA libraries, expressed sequence tags (ESTs), and a cDNA microarray for the study of insect-induced defenses in poplar Molecular Ecology 15: 1275-1297
Thompson, SL and Ritland, K (2006). A novel mating system analysis for modes of self-oriented mating applied to diploid and polyploid arctic Easter daisies (Townsendia hookeri) Heredity 97: 119-126
Ally D, Ritland K (2006). A case Study: looking at the effects of fragmentation on genetic structure in different life history stages of old-growth mountain hemlock (Tsuga mertensiana) J. Heredity 98: 73-78
Tuskan et al. (including K. Ritland) (2006). The genome of black cottonwood, Populus trichocarpa (Torr. & Gray) Science 313: 1596-1604
Goodwille, C., Ritland, C., and Ritland, K. (2006). The genetic basis of floral traits associated with mating system evolution in Leptosiphon (Polemoniaceae): an analysis of quantitative trait loci Evolution 60: 491-504
Steven Ralph, Hesther Yueh, Michael Friedmann, Dana Aeschliman, Jeffrey A. Zeznik, Colleen C. Nelson, Yaron S. N. Butterfield, Robert Kirkpatrick, Jerry Liu, Steven J. M. Jones, Marco A. Marra, Carl J. Douglas, Kermit Ritland and Jörg Bohlmann (2006). Conifer Defense against Insects: Microarray Gene Expression Profiling of Sitka Spruce (Picea sitchensis) Induced by Mechanical Wounding or feeding by Spruce Budworms (Choristoneura occidentalis) or White Pine Weevils (Pissodes strobi) Plant, Cell and Environment
Ehlting, J., Mattheus, N., Aeschliman, D. S., Li, E., Hamberger, B., Cullis, I.F, Zhuang, J., Kaneda, M., Mansfield, S. D., Samuels, A. L., Ritland, K., Ellis, B. E., Bohlmann, J. Douglas, C. J. (2005). Global transcript profiling of primary stems from Arabidopsis thaliana identifies candidate genes for missing links in lignin biosynthesis and transcriptional regulators of fiber differentiation The Plant Journal. 42: 618-640
Ritland K, Ralph S, Lippert, D, Rungis D, Bohlmann J (2005). A new direction for conifer genomics Kluwer-Springer Press Pages 75-84 in” Landscapes, Genomics and Transgenic Conifer Forests” ed. Claire G. Williams, 261 pp.
Ritland K (2005). Multilocus estimation of pairwise relatedness with dominant markers Molecular Ecology 14: 3157-3165
Beland JD, Krakowski J, Ritland C, Ritland K, El-Kassaby YA (2005). Genetic structure and mating system of northern Arbutus menziesii populations Can. J. Botany 83: 1581-1589
Rungis D, Hamberger B, Bérubé Y, Wilkin J, Bohlmann J, Ritland, K (2005). Efficient genetic mapping of nucleotide polymorphisms based upon DNA mismatch digestion Molecular Breeding 16: 261-270.
Wilkin JE, Shamoun SF, Ritland C, Ritland K, El-Kassaby, YA (2005). Population Genetics of Gaultheria shallon in British Columbia and the Implications for Management using Biocontrol Can. J. Botany 83: 501-509
van Kleunen, M. and Ritland, K (2005). Estimating heritabilities and genetic correlations with marker-based methods: an experimental test in Mimulus guttatus J. Heredity 96: 368-375.
Lippert D, Zhuang J, Ralph S, Ellis D, Gilbert M, Olafson R, Ritland K, Ellis B, Douglas C, Bohlmann J (2005). Proteome analysis of early somatic embryogenesis in Picea glauca Proteomics 5: 461-473
O’Connell L, Russel J, Ritland K (2005). Post-pollination mechanisms promoting outcrossing in a self-fertile conifer Can. J. Botany 83: 335-342
Wahbe TR, Ritland C, Bunnell FL, Ritland K (2005). Population genetic structure of tailed frogs (Ascaphus truei) in clearcut and old-growth stream habitats in south coastal British Columbia Can J Zoology 83: 1460-1468
Kolosova, N., Miller, B., Ralph, S., Ellis, B.E., Douglas, C., Ritland, K., Bohlmann, J. (2004). Improved Protocol for Isolation of High-Quality mRNA from Conifers and Poplars. BioTechniques 36: 821-824
van Kleunen, M. and Ritland, K. (2004). Predicting evolution of floral traits associated with mating system in a natural plant population J. Evol. Biol. 17: 1389-1399
O’Connell, L. M. and K. Ritland. (2004). Somatic mutation rate at microsatellite loci in Thuja plicata. Journal of Heredity 95: 172-176
Wellman, H., Ritland, C. and Ritland, K. (2004). Genetic effects of domestication in western hemlock, Tsuga heterophylla. Forest Genetics 10: 229-240
Ritland, K. (2004). Pathways to plant population genomics. NRC Research Press, Ottawa, Ontario. In Molecular Genetics and Ecology of Plant Adaptation. Proceedings of an International Workshop held December 11–13, 2002, in Vancouver, British Columbia, Canada. Edited by Q.C.B. Cronk, J. Whitton, R.H. Ree, and I.E.P. Taylor. pp. 11–17.
Ritland, K and Travis, S. (2004). Inferences involving individual coefficients of relatedness and inbreeding in natural populations of Abies. Forest Ecology and Management 197: 171-180
Travis, S. E., Proffitt, C.E., Ritland, K. (2004). The successional dynamics of fine-scale genetic structure and its effects on inbreeding in a facultatively clonal plant (Spartina alterniflora). Ecological Applications 14: 1189-1202
Iddrisu, M. and Ritland, K. (2004). Genetic variation, population structure and mating system in bigleaf maple (Acer macrophyllum Pursh) Can. J. Botany 82: 1817-1825
Ritland, K. and LeBlanc, M. (2004). Mating system of four inbreeding monkeyflower (Mimulus) species as revealed by “progeny-pair” analysis of highly informative microsatellite markers. Plant Species Biology 19: 149-157
O’Connell, L., Russel, J. and Ritland, K. (2004). Fine-scale estimation of outcrossing in western redcedar with microsatellite assay of bulked DNA. Heredity 93: 443-449
Hu, X-S., Goodwillie, C. and Ritland, K. (2004). Joining genetic linkage maps using a joint likelihood function. Theor. Appl. Genetics 109: 996-1004
Rungis, D., Bérubé, Y., Zhang, J., Ralph, S., Ritland, C.E., Ellis, B.E., Douglas, C., Bohlmann, J. Ritland, K. (2004). Robust simple sequence repeat (SSR) markers for spruce (Picea spp.) from expressed sequence tags (ESTs). Theor. Appl. Genet. 109: 1283-1294
Bérubé, Y, Ritland, CE and Ritland, K. (2003). Isolation, characterization, and cross-species utility of microsatellites in yellow cedar, (Chamaecyparis nootkatensis) Genome 46: 353-361.
Ritland, K. (2002). Estimation of of gene frequency and heterozygosity from pooled samples. Molecular Ecology Notes 78. 2: 370-372.
Marshall, H.D. and Ritland, K. (2002). Genetic diversity and differentiation of Kermode bear populations Molecular Ecology 11: 685-697
Ritland, K. (2002). Extensions of models for the estimation of mating systems using n independent loci Heredity 88: 221-228
Marshall, H.D., Newton, C., and Ritland, K (2002). Chloroplast phylogeography and the evolution of highly polymorphic microsatellites in Lodgepole pine (Pinus contorta). Theoretical and Applied Genetics 104: 367-378
Marshall, H.D., Newton, C. and Ritland, K. (2001). Sequence-repeat polymorphisms exhibit the signiture of recombination in lodgepole pine. Mol. Biol. Evol. 11: 2136-2138.
Ritland, K., Newton, C., and Marshall, D. (2001). Inheritance and population structure of the white-phased “Kermode” black bear. Current Biology 11: 1468-1472.
Klaper, R.D., K. Ritland, T.A. Mousseau, and M.D. Hunter. (2001). Heritability of phenolics in a natural population of Quercus laevis using molecular markers. J. Heredity 92: 421-426.
Ritland, C., T. Pape and K. Ritland. (2001). Genetic structure of yellow-cedar (Chamaecyparis nootkatensis). Canadian Journal of Botany 79: 822-828.
O’Connell, L., F. Viard and K. Ritland. (2001). The mating system in natural populations of western red cedar (Thuja plicata). Canadian Journal of Botany 79:753-756.
Ally, D., K. Ritland and Y.A. El-Kassaby. (2000). Genetic diversity, differentiation and mating system in mountain hemlock (Tsuga mertensiana) across British Columbia. For. Genetics 7:97-108.
Ritland, Carol E. and Kermit M. Ritland. (2000). DNA fragment markers in plants. Blackwell Scientific, Oxford, U.K. Pp. 208-234 in Allan J. Baker (ed.), Molecular Methods in Ecology.
Ritland, K. (2000). Marker-inferred relatedness as a tool for detecting heritability in nature. Molecular Ecol. 9:1195-1204.
Ritland, K. (2000). Detecting inheritance with inferred relatedness in nature. Oxford University Press. Pp. 187-199 in T.A. Mousseau, B. Sinervo and J. Endler (eds.), Adaptive Genetic Variation in the Wild.
Ritland, K., L.A. Dupuis, F.L. Bunnell, W.L.Y. Hung, and J.E. Carlson. (2000). Phylogeography of the tailed frog (Ascaphus truei) in British Columbia. Can. J. Zoology 78:1749-1758.
Lynch, M. and K. Ritland. (1999). Estimation of pairwise relatedness with molecular markers. Genetics. 152: 1753-1766.
Mousseau, T.A., K. Ritland and D.D. Heath. (1998). A novel method for estimating heritability using molecular markers. Heredity 80:218-224.
Sun, M. and Ritland, K. (1998). Mating system parameters of yellow starthistle (Centaurea solstitialis), a successful colonizer in North America. Heredity 80:225-232.
Travis, S.E., K. Ritland, T.G. Whitham and P. Keim (1998). A genetic linkage map of pinyon pine (Pinus edulis) based on amplified fragment length polymorphisms. Theor. Appl. Genetics 97: 871-880.
Awadalla, P. and Ritland, K. (1997). Microsatellite variation in Mimulus species of contrasting mating systems. Molecular Biology and Evolution 14: 1023-1034.
Lin, J.-Z. and Ritland, K. (1997). Quantitative trait loci differentiating the outbreeding Mimulus guttatus from the inbreeding Mimulus platycalyx. Genetics 146:1115-1121.
Fu, Y.B and K. Ritland. (1996). Marker-based inferences about epistasis for genes influencing inbreeding depression. Genetics 144:339-348.
Lin, J. and K. Ritland. (1996). The effects of selective genotyping on estimates of proportion of recombination between linked quantitative trait loci. Theoretical and Applied Genetics 93:1261-1266.
Lin, J. and Ritland, K. (1996). Construction of a genetic linkage map in the wild plant Mimulus using RAPD and isozyme markers. Genome 39:63-70.
Ritland, K. (1996). Inferring the genetic basis of inbreeding depression in plants. Genome 39:1-8.
Ritland, K. (1996). Estimators for pairwise relatedness and individual inbreeding coefficients. Genetical Research 67:175-185.
Ritland, K. (1996). A marker-based method for inferences about quantitative inheritance in natural populations. Evolution 50:1062-1073.
Ritland, K. , Y.B. Fu and J.Z. Lin. (1996). Characterizing molecular quantitative variability for inbreeding depression. Plant Species Biology 11:23-31.
Ritland, K. and C. Ritland. (1996). Inferences about quantitative inheritance based upon natural population structure in the common monkeyflower, Mimulus guttatus. Evolution 50: 1074-1082.