Forest Sciences Centre 3041
2424 Main Mall
Vancouver, BC V6T 1Z4
phone: 6048229695

Areas of research include:

  • Ecology, life history variation, behaviour and conservation of birds in high elevation habitats
  • Metapopulation and connectivity processes for alpine bird populations
  • Avian use of mountain habitats during migration
  • Nest Webs – cavity nesting bird community dynamics in natural and managed forests in relation to forest cutting regimes and natural disturbance events
  • The roles of excavating birds in cavity nesting community dynamics
  • Effectiveness of avian monitoring programs, surrogates and indicator species


Ecology and Life History of Mountain Birds


Life history variation of vertebrates across elevations


Effects of forest management and forest health on avian biodiversity of mixed forest ecosystems in Canada and Chile

Environment Canada

Structure and function of cavity-nester communities in the Americas

NSERC and Environment Canada

Professional Affiliations

Advisory Commitee for UBC Research Forests
Bird Studies Canada Scientific Advisory Board, Scientific Advice, January 2015
British Ornithological Society; Member
Canadian Society for Ecology and Evolution CSEE, Member, January 2016
Cooper Ornithological Society; Member, Member, January 2015
International Ornithological Union IOC, Scientific Program Committee, January 2015
IUCN/ICBP (The World Conservation Union) Specialist Group for Galliformes; Canadian Representative, 1996-current, Member, January 2016
Natural Sciences and Engineering Research Council NSERC, Reviewer, January 2014
NSERC Grant Selection Committee 18 -Evolution and Ecology; Member, 2001-04
The Wildlife Society Member, Member , January 2016
University of Alberta, Challenge for Biodiversity Awards; Referee, 1995-current, Reviewer, January 2014
Wilson Ornithological Society; Member, Member, January 2017
Society of Canadian Ornithologists;, Member, January 2017
American Ornithological Society , President-Elect, August 2016


Doris Huestis Speirs Award, Outstanding Lifetime Contributions to Canadian Ornithology, Canada , 2008
5NR Science Award to Leaders in Sustainable Development, Natural Resources, Canada , 2003

Current Graduate Students

Devin deZwaan, PhD
Nestling development strategies in horned larks breeding in northern alpine habitats
Jennifer Greenwood, Phd
Alpine songbird life history and coping mechanisms
Julien StAmand, MSc
Critical Habitat Attributes for the red-listed Williamson’s Sapsucker in British Columbia
Jennifer Nielsen, MSc
Forest Biodiversity and Habitat Attributes

Post Doctoral Fellows and Research Assistants

Dr Tomás (José) Ibarra
Dr. Anna Drake
Dr. Michelle Jackson

Current Courses

Selected Publications

Wesolowski, T. and K. Martin (2017). Tree holes and hole nesting birds in European and North American Forests, In: Ecology and Conservation of Forest Birds. Mikusinski, G, J-M Roberge, and R.J. Fuller, Eds. Ecology, Biodiversity and Conservation Series, Cambridge Univ Press, UK.
Cockle, K. L., A. Bodrati and K. Martin (2017). Persistence of nesting cavities in the subtropical Atlantic Forest: influence of habitat, tree and cavity characteristics Forest Ecology and Management, 384: 200-207
de Zwaan, D.R., J. L. Greenwood and K. Martin (2017). Feather melanin and micro-structure variation in Dark-eyed Junco (Junco hyemalis) across an elevational gradient in the Selkirk Mountains Journal of Avian Biology, In press
Ibarra, J. T., T.A Altamirano, A. Vermehren, F. Hernán Vargas, and K. Martin (2017). Observations of a tree-cavity nest of the rufous–legged owl and predation of an owl nestling by a chimango caracara in Andean temperate forests J. Raptor Research, 51: 85-88. doi/full/10.3356/JRR-16-68.1
Martin, K., S. D. Wilson, E.C. MacDonald, A. F. Camfield, M. Martin, and S. A. Trefrey (2017). Effects of severe weather on reproduction for sympatric songbirds in an alpine environment: interactions of climate extremes influence nesting success The Auk, in press
Goullaud, E.L., D. de Zwaan, and K. Martin (2017). Predation risk-induced adjustments in provisioning behavior for horned larks (Eremophila alpestris) in British Columbia Wilson Journal of Ornithology,
Honorato, M. T., T. A. Altamirano, J. T. Ibarra, M. De la Maza, C. Bonacic, K. Martin (2016). Composition and preference for nesting materials of cavity nester vertebrates in temperate Andean forests of Chile Revista Bosque, 37: 485-492. DOI: 10.4067/S0717-92002016000300005 [in Spanish].
Beckmann, C, and K. Martin. (2016). Testing hypotheses about the function of repeated nest abandonment as a life history strategy in a passerine bird. Ibis, 158: 335-342, DOI: 10.1111/ibi.12361
Jackson, M. M., E. Topp, S. E. Gergel, K. Martin, F. Pirotti, T. Sitzia. (2016). Expansion of subalpine woody vegetation over 40 years on Vancouver Island, British Columbia, Canada. Canadian J. Forest Research, 46: 437-443, doi 10.1139/cjfr-2015-0186
Huang A. C., J. E. Elliott, S. Hindmarch, S. Lee, F. Maisonneuve, V. Bowes, K. M. Cheng, and K. Martin (2016). Increased rodenticide exposure rate and risk of toxicosis in barn owls (Tyto alba) from southwestern Canada and linkage with demographic but not genetic factors. Ecotoxicology, 25: 1061-1071. DOI 10.1007/s10646-016-1662-6
Boyle, W.A., B. K. Sandercock, K. Martin (2016). Patterns and drivers of intraspecific variation in avian life history along elevational gradients: a meta-analysis. Biological Reviews. 91: 469-482. DOI: 10.1111/brv.12180
Seneviratne, S. S., P. Davidson, K. Martin and D. Irwin (2016). Low levels of hybridization across two contact zones among three species of woodpeckers (Sphyrapicus sapsuckers) Journal of Avian Biology. 47: DOI 10.1111/jav.00946
Huang, A., J. E. Elliott, K. M. Cheng, K. Ritland, C. Ritland, S. K. Thompson, S. Hindmarch, and K. Martin (2016). Barn owls (Tyto alba) in North America: Phylogeographic structure, connectivity, and genetic diversity. Conservation Genetics 17: 357-367
Floyd, C., and K. Martin (2016). Avian Ecosystem Engineers: Birds that excavate cavities. Chapter 10,pp 296-318. In: Why Birds Matter, C. Skekercioglu, D. G. Wenny, C. J. Whelan (editors). University of Chicago Press.
Hunter, M. Jr., P. Barton, A. Calhoun, J. Pierson, A. Tulloch, M. Westgate, M. Beger, C. Branquinho, T. Caro, J. Gross, J.Heino, P. Lane, C. Longo, K. Martin, W. H. McDowell, C. Mellin, H. Salo, D. Lindenmayer (2016). Two roles for ecological surrogacy: indicator surrogates and management surrogates. Ecological Indicators, 63: 121-125
MacDonald, E.C., A. F. Camfield, M. Martin, S. Wilson, and K. Martin (2016). Nest-site selection and consequences for nest survival among three sympatric songbirds in an alpine environment. Springer Journal of Ornithology, 157: 393-405. DOI 10.1007/s10336-015-1286-2
Drever, M.C., L.W. Gyug, j. Nielsen, A.K. Stuart-Smith, I.P. Ohanjanian, K.T. Fort, K. Martin (2015). Using field data and GIS-derived variables to model occurrence of Williamson’s Sapsucker nesting habitat at multiple spatial scales. PLOS ONE doi:10.1371/journal.pone.0130849. 10(7): e0130849
Martin, K., L. A. Robb, S. D. Wilson and C. E. Braun (2015). White-tailed Ptarmigan. In the Birds of North America, No. 68 (A. Poole, Ed.). Cornell Lab of Ornithology The Academy of Natural Sciences; Washington, DC: pp. 1-28.
Boyle, W.A., and K. Martin. (2015). The conservation value of high elevation habitats to North American migrant birds. Biological Conservation, 192: 461-476.
Jackson, M. M., S. E. Gergel, and K. Martin. (2015). Effects of climate change on habitat availability and configuration for an endemic coastal bird. PLoS ONE 10(11): e0142110. doi:10.1371/journal. pone.0142110
Ibarra, J. T., and K. Martin. (2015). Biotic homogenization: loss of avian functional richness and habitat specialists in disturbed Andean temperate forests. Biological Conservation, 192: 418-427.
Lindenmayer, D, J. Pierson, P. Barton, M. Beger, C. Branquinho, A. Calhoun, T. Caro, H. Greig, J. Gross, J. Heino, M. Hunter, P. Lane, C. Longo, K. Martin, W. H. McDowell, C. Mellin, H. Salo, A. Tulloch, and M. Westgate. (2015). A new framework for selecting environmental surrogates. Total Science of the Environment, 538: 1029-1038. DOI: 10.1016/j.scitotenv.2015.08.056
Martin, K. J. T., Ibarra, and M. Drever. (2015). Avian surrogates in terrestrial ecosystems: theory and practice. Chapter 5, Pp 33-44 In: Indicators and Surrogates in Ecology, Biodiversity and Environmental Management, Edited by D. Lindenmayer, P. Barton and J. Pierson. CSIRO Publishing, Melbourne. CRC Press, London.
Ibarra, J. T., and K. Martin (2015). Beyond species richness: an empirical test of top predators as surrogates for functional diversity and endemism Ecosphere 6 (8): 142.
Beckmann, C., P. A. Biro, and K. Martin (2015). Hierarchical analysis of avian re-nesting behavior: mean, across-individual, and intra-individual responses. Behavioral Ecology and Sociobiology 69:1631–1638. DOI 10.1007/s00265-015-1974-1
Cockle, K. L. and, K. Martin. (2015). Temporal dynamics of a commensal network of cavity-nesting vertebrates: increased diversity during an insect outbreak. Ecology. 96: 1093-1104.
Cockle, K. C., A. Bodrati, M. Lammertink, and K. Martin. (2015). Cavity characteristics, but not habitat, influence nest survival of cavity-nesting birds along a gradient of human impact in the subtropical Atlantic forest. Biological Conservation, 184:193-200. DOI: 10.1016/j.biocon.2015.01.026
Jackson, M., S. E. Gergel and K. Martin. (2015). Citizen science and field survey observations provide comparable results for mapping Vancouver Island White-tailed Ptarmigan (Lagopus leucura saxatilis) distribution. Biological Conservation, 181: 162-172. DOI:10.1016/j.biocon.2014.11.010
Ibarra, J. T., T. A. Altamirano, G. Vergara, F. H. Vargas, and K. Martin. (2015). Natural history and autecology of the Austral Pygmy-Owl (Glaucidium nana) in the Andean temperate forest of the Araucania region, southern Chile. In Edicion Especial: Aves Rapaces Del Bosque Templado Austral. Boletín Chileno de Ornitología 21: 29−40
Martin, K. (2015). Nest webs and woodpecker ecological services: the role of woodpeckers in tree cavity-using wildlife communities in North America. In: Woodpeckers – New Insights Into Their Biology, H. Winkler, ed. Biologie Zentrum, Denisia 36, zugleich Kataloge des oberösterreichischen Landesmuseums Neue Serie 164: 77-86
MacDonald, E.C. A. F. Camfield, Jill E. Jankowski, and K. Martin (2014). An alpine breeding songbird can adjust dawn incubation rhythms to annual thermal regimes. The Auk, 131:495-506. DOI: 10.1642/AUK-13-234.1
Ibarra, J. T., K. Martin , T. A. Altamirano, F. H. Vargas, C. Bonacic (2014). Factors associated with the detectability of Owls in South–American Temperate Forests: Implications for Nocturnal Raptor Monitoring. Journal of Wildlife Management, 78:1078-1086. DOI: 10.1002/jwmg.740
Edworthy, A. B. and K. Martin. (2014). Long-term dynamics of the characteristics of tree cavities used for nesting by vertebrates. Forest Ecology and Management, 334: 122-128.
Ibarra, J. T., K. Martin, M. C. Drever, and G. Vergara. (2014). Occurrence patterns and niche relationships of sympatric owls in South American temperate forests: a multi-scale approach. Forest Ecology and Management, 331: 281-291. http:
Martin, K. (2014). Avian Strategies for living at high elevation: life-history variation and coping mechanisms in mountain habitats. IBIS-Conference Proceeding. http//www.
Norris, A.R., and K. Martin (2014). Direct and indirect effects of an insect outbreak increase the reproductive output for an avian insectivore and nest-cavity excavator, the red-breasted nuthatch (Sitta canadensis). Journal of Avian Biology, 45:280-290. DOI: 10.1111/jav.00112
Filotas, E., Parrott, L., Burton, P.J., Chazdon, R.L., Coates, K.D., Coll, L., Haeussler, S., Martin, K., Nocentini, S., Puettmann, K.J., Putz, F.E., Simard, S.W., and Messier, C. (2014). Viewing forests through the lens of complex systems science. ESA Ecosphere, 5:art1, pp1-23.
Robles, H and K. Martin (2014). Habitat-mediated variation in the importance of ecosystem engineers for secondary cavity nesters in a nest web. PLoS ONE 9(2): e90071. DOI: 10.1371/journal.pone.0090071
Ibarra, J. T., T. A. Altamirano, K. Martin, F. H. Vargas, C. Bonacic. (2014). Tree cavity-nesting of Austral Pygmy Owls (Glaucidium nana) in Andean Temperate Forests of southern Chile. J. Raptor Research, 48: 82-85.
Krebs, C.J., R. Boonstra, S. Boutin, A.R.E. Sinclair, J.N.M. Smith, B.S. Gilbert, K. Martin, M. O’Donoghue, and R. Turkington (2014). Trophic Dynamics of the Boreal Forests of the Kluane Region. Arctic, 67: Supplement 71-81.
Edworthy, A.B., and K. Martin. (2013). Persistence of tree cavities used by cavity-nesting vertebrates declines in harvested forests. Journal of Wildlife Management, 77: 770-776. DOI: 10.1002/jwmg.526
MacDonald, E.C., A.F. Camfield, J.E. Jankowski, and K. Martin (2013). Extended incubation recesses by alpine-breeding Horned Larks: a strategy for dealing with inclement weather? Association of Field Ornithologists Journal of Field Ornithology, 84: 58-68. DOI: 10.1111/jofo.12006
Simard, S.W., Martin, K., Vyse, A., and Larson, B. (2013). Meta-networks of fungi, fauna and flora as agents of complex adaptive systems: an example from the interior Douglas-fir forests of southern British Columbia. Chapter 7, pages 133-164. In: Managing World Forests as Complex Adaptive Systems: Building Resilience to the Challenge of Global Change. Edited by Puettmann, K, Messier, C, and Coates, KD. Routledge, NY. ISBN 978-0-415-51977. 369pp.
K. Martin (2013). The Ecological Values of Mountain Environments and Wildlife. Bentham Science Publishers Pp. 3-29, Chapter 1 in Rixen and Rolando (eds.), The Impact of Skiing on Mountain Environments.
Robles, H and K. Martin (2013). Resource quantity and quality determine the inter-specific associations between ecosystem engineers and resource users in a cavity-nest web. PLOS ONE, 8(9): e74694. DOI:10.1371/journal.pone.0074694
Evans Ogden, L. J., K. Martin, and T. D. Williams. (2013). Elevational differences in estimated fattening rates suggest high-elevation sites are high quality habitats for fall migration. The Auk, 130:98-106.
Norris, A.R., M.C. Drever, K. Martin. (2013). Insect outbreaks increase populations and facilitate reproduction in a cavity-dependent songbird, the Mountain Chickadee (Poecile gambeli). Ibis, 155: 165-176. DOI: 10.1111/ibi.12005
Aitken, K.E.H., and K. Martin. (2012). Experimental test of nest-site limitation in mature mixed forests of central British Columbia, Canada. Journal of Wildlife Management, 76: 557-565. DOI:10.1002/jwmg.286
Zhu, X., D. Srivastava, J.N.M. Smith, and K. Martin (2012). Habitat selection and reproductive success of Lewis' Woodpecker (Melanerpes lewis) at its northern limit. PLOS ONE 7 (9): e44346, pp. 1-9. DOI: 10.1371/journal.pone.0044346
Edworthy, A.B., K.L. Wiebe and K. Martin. (2012). Survival analysis of a critical resource for cavity-nesting communities: patterns of tree cavity longevity. Ecological Applications, 22: 1733-1742.
Wilson, S., and K. Martin. (2012). Influence of life history strategies on sensitivity, population growth and response to climate for sympatric alpine birds. BMC Ecology,12:9. DOI:10.1186/1472-6785-12-9
Norris, A.R., and K. Martin. (2012). Red-breasted nuthatches (Sitta canadensis;) increase cavity excavation in response to a mountain pine beetle (Dendroctonus ponderosae) outbreak. Ecoscience, 19: 308-315. DOI: 10.2980/19-4-3519
Cockle, K., K. Martin, and G. Robleda. (2012). Linking fungi, trees, and hole-using birds in a Neotropical tree-cavity network: pathways of cavity production and implications for conservation. Forest Ecology and Management 264: 210-219. DOI: 10.1016/j.foreco.2011.10.015
Koch, A.J., K. Martin, and K.E.H. Aitken. (2012). The relationship between introduced European Starlings and the reproductive activities of Mountain Bluebirds and Tree Swallows in British Columbia, Canada. Ibis,154: 590-600. DOI: 10.1111/j.1474-919X.2012.01242.x
Evans Ogden, L.J., M. Martin, and K. Martin. (2012). Mating and breeding success decline with elevation for the Pacific wren (Troglodytes pacificus) in coastal mountain forests. Wilson Journal of Ornithology, 124: 270-276. DOI:
Blanc, L.A., and K. Martin. (2012). Identifying suitable woodpecker nest trees using decay selection profiles in trembling aspen (Populus tremuloides). Forest Ecology and Management, 286: 192-202. DOI: 10.1016/j.foreco.2012.08.021
Koch, A., M.C. Drever, K. Martin. (2011). The efficacy of common species as indicators; avian responses to disturbance in British Columbia, Canada. Biodiversity and Conservation, 20: 3555-3575. DOI: 10.1007/s10531-011-0148-3
Cockle, K.L., K. Martin, T. Wesolowski (2011). Woodpeckers, decay and the future of cavity-nesting vertebrate communities worldwide. Ecological Society of America Frontiers in Ecology and the Environment, 9: 377-382, doi: 10:1890/110013
Camfield, A.F., S.F. Pearson, and K. Martin. (2011). A demographic model to evaluate population declines in endangered Streaked Horned Lark. Avian Conservation and Ecology 6 (2): 4.
Fedy, B.C., and K. Martin. (2011). The influence of fine-scale habitat features on regional variation in population performance of alpine white-tailed ptarmigan. Condor 113: 306-315.
Sandercock, B.K., K. Martin, and G. Segelbacher (eds.) (2011). Ecology, Conservation and Management of Grouse. Studies in Avian Biology, No. 39. University of California Press, Berkeley, CA.
Martin, K., and S. Wilson (2011). Ptarmigan in North America: Influence of life history and environmental conditions on population persistence -in R. T. Watson, T. J. Cade, M. Fuller, G. Hunt, and E. Potapov (Eds.). Gyrfalcons and ptarmigan in a changing world. Vol 1, pp 45-54. The Peregrine Fund, Boise, Idaho, USA. DOI 10.4080/gpcw.2011.0105
Edworthy, A., M.C. Drever, K. Martin (2011). Woodpeckers increase in abundance but maintain fecundity in response to an outbreak of mountain pine bark beetles. Forest Ecology and Management. 261: 203-210. DOI:10.1016/j.foreco.2010.10.006
Wilson, S., and K. Martin. (2011). Life history and demographic variation in an alpine specialist at the latitudinal extremes of the range. Population Ecology 53: 459–471. DOI: 10.1007/S10144-011-0261-X
Martin, K., S. Wilson and S. J. Hannon (2011). Mechanisms underlying variation in renesting ability of willow ptarmigan. Pp. 233–246 in B. K. Sandercock, K. Martin, and G. Segelbacher (editors). Ecology, conservation, and management of grouse. Studies in Avian Biology (no. 39), University of California Press, Berkeley, CA.
Wilson, S., and K. Martin. (2010). Variable reproductive effort for two sympatric ptarmigan in response to spring weather conditions in a northern alpine ecosystem. Journal of Avian Biology, 41:319-326. DOI: 10.1111/j.1600-048X.2009.04945.x
Camfield, A .F., S. F. Pearson, and K. Martin. (2010). Life history variation between high and low elevation subspecies of horned larks. Journal of Avian Biology, 41: 273-281. DOI:10.1111/j.1600-048X.2009.04816.x
Drever, M.C.,and K. Martin. (2010). Response of woodpeckers to changes in forest health and harvest: implications for conservation of avian biodiversity. Forest Ecology Management, 259: 958-966. DOI:10.1016/j.foreco.2009.11.038
Norris, A.R., and K. Martin. (2010). The perils of plasticity: Dual resource pulses increase facilitation but destabilize populations of small cavity-nesting vertebrates. Oikos, 119: 1126-1135. DOI: 10.1111/j.1600-0706.2009.18122.x
Norris, A, K. Cockle, and K. Martin (2010). Evidence for tolerance of parasitism in a tropical cavity-nesting bird, the planalto woodcreeper (Dendrocolaptes platyrostris), in northern Argentina Cambridge University Press – Journal of Tropical Ecology, 26: 619-626
Cockle, Kristina, K. Martin and Mark C. Drever (2010). Supply of tree-holes limits nest density of cavity-nesting birds in primary and logged subtropical Atlantic Forest Biological Conservation, 143: 2851-2857
Wong, M.L., B.C. Fedy, S. Wilson, and K. Martin. (2009). Adoption in Rock and White-tailed Ptarmigan. The Wilson Journal of Ornithology. 121: 638-641
Camfield, A .F., and K. Martin. (2009). The influence of ambient temperature on horned lark incubation behaviour in an alpine environment. Behaviour, 146:1615-1633.
Martin, M., A.F. Camfield, and K. Martin. (2009). Demography of an alpine population of Savannah Sparrows. Journal of Field Ornithology. 80: 253-264.
Bears, H., K. Martin, and G.C. White. (2009). Breeding in high-elevation habitat results in shift to slower life-history strategy within a single species. Journal of Animal Ecology 78:365-375.
Drever, M.C., J. R. Goheen, and K. Martin (2009). Species-energy theory, pulsed resources, and the regulation of avian richness during a mountain pine beetle outbreak. Ecology 90:1095-1105.
Holmes, D.J. and K. Martin (2009). A bird’s eye view of aging: What’s in it for Ornithologists? Auk 126:1-23.
Mahon, C.L., J.D. Steventon, and K. Martin (2008). Cavity and bark nesting bird responses to partial cutting in northern conifer forests. Forest Ecology and Management, 265: 2145-2153.
Nouvet, S., S. Wilson and K. Martin (2008). Confirmed breeding of four shorebird species, the Surfbird, Upland Sandpiper, American Golden Plover, and Wandering Tattler, in the Ruby Mountains of the southwest Yukon Territory Western birds 39: 22-30.
Mahon, C.L, K. Martin, and V. LeMay (2008). Do cross-scale correlations confound analysis of nest site selection for chestnut-backed chicadees? Cooper Ornithological Society The Condor 110: 563-568
Rangel-Salazar, J.L., K. Martin, P.L. Marshall and R.W. Elner. (2008). Population dynamics of the Ruddy-capped Nightingale Thrush (Catharus frantzii) in Chiapas, Mexico: influences of density, productivity and survival. Journal of Tropical Ecology 24: 583-593
Cockle, K., K. Martin, and K. L. Wiebe (2008). Availability of cavities for nesting birds in the Atlantic Forest, Argentina. Ornitologia Neotropical 19(Suppl.):269-278.
Aitken, K. E. H., and K. Martin (2008). Resource selection plasticity and community responses to experimental reduction of a critical resource. Ecology 89: 971-980.
Fedy, B. C., K. Martin, C. Ritland, and J. Young. (2008). Genetic and ecological data provide incongruent interpretations of population structure and dispersal in naturally subdivided populations of white-tailed ptarmigan (Lagopus leucura). Molecular Ecology 17: 1905-1917.
Drever, M. C., K. E. H. Aitken, A. R. Norris, and K. Martin. (2008). Woodpeckers as reliable indicators of bird richness, forest health and harvest. Biological Conservation 141: 624-634.
Cornelius, C., K. Cockle, N. Politi, I. Berkunsky, L. Sandoval, V. Ojeda, L. Rivera, Jr. M. Hunter, and K. Martin (2008). Cavity-nesting birds in neotropical forests: Cavities as a potentially limiting resource. Ornitologia Neotropical 19(Suppl.): 253-268.
Wilson, S. and K. Martin (2008). Breeding habitat selection of sympatric rock, white-tailed, rock and willow ptarmigan in the southern Yukon Territory, Canada. Journal of Ornithology. 149: 629-637.
Rangel-Salazar, J.L., K. Martin, P. Marshall, and R.W. Elner. (2008). Habitat variation, nest-site selection and parental behaviour influences on breeding success of the ruddy-capped nightingale thrush (Catharus frantzii) in Chiapas, Mexico Auk 125: 358-367.
Bears, H., M.C. Drever and K. Martin (2008). Comparative morphology of dark-eyed juncos breeding at two elevations: a common aviary experiment Journal of Avian Biology 39: 152-162.
Norris, A. R. and K. Martin (2008). Mountain pine beetle presence affects nest patch choice of red-breasted nuthatches. Journal of Wildlife Management 72: 733-737.
Drever, M.C. and K. Martin (2007). Spending time in the forest: responses of cavity nesters to temporal changes in forest health in interior British Columbia. Springer-Verlag, New York. Chapter 13. Pp. 236-251 in J. Bissonnette and I. Storch (eds), Temporal explicitness in landscape ecology; Understanding wildlife responses to changes in time.
Martin, K. and A.R. Norris (2007). Life in the small-bodied cavity nester guild: Demography and reproduction of sympatric Mountain and Black-capped Chickadees within Nest Web communities under changing habitat conditions. Oxford University Press, UK. Chapter 8. Pp. 111-130 in Ken Otter (ed.), Ecology and behavior of chickadees and titmice; An integrated approach.
Mennill, D.J., T.M. Burg, R.L. Curry, K. Martin, A.R. Norris, L. Ratcliffe, M.W. Reudink, L.M. Rossano, and K. A. Shubert (2007). Parid reproductive behavior. Oxford University Press, UK. Chapter 9. Pp. 147-150 in Ken Otter (ed.), Ecology and behavior of chickadees and titmice; An integrated approach.
Mahon, C.L., K. Martin, and J.D. Steventon (2007). Habitat attributes and chestnut-backed chickadee nest site selection in uncut and partial cut forests Canadian Journal of Forest Research 37(7): 1272-1285.
Camfield, A.F., A.J. Clason, and K. Martin (2007). Mother-son parental care in horned larks Wilson Journal of Ornithology 199: 302-304.
Wiebe, K.L., W.D. Koenig, and K. Martin (2007). Costs and benefits of nest reuse versus excavation in cavity-nesting birds Annales Zoologica Fennici 44: 209-217.
Aitken, K.E.H. and K. Martin (2007). The importance of excavators in hole nesting communities: availability and use of natural tree holes in old and mixed forests of western Canada Journal of Ornithology 148 (Suppl. 2): 425-434.
Squires, K.A., K. Martin and R.I. Goudie (2007). Vigilance behavior in the harlequin duck (Histrionicus histrionicus) during the pre-incubation period in Labrador. The Auk 124:241-252.
Topp, S., L.J. Evans Ogden, and K. Martin (2007). Apparent absence of blood parasites in winter wrens in British Columbia. Journal of Field Ornithology 78: 308-313.
Wilson. S, K. Martin, S.J. Hannon (2007). Nest survival patterns in willow ptarmigan: influence of time, nesting stage and female characteristics Condor 109:377-388.
Martin, K. and K.L. Wiebe (2006). Impacts of extreme climate events on fitness of alpine birds Acta Zoologica Sinica 52 (Supplement):162-164.
Mahon, C.L. and K. Martin (2006). Nest survival of chickadees in managed forests: habitat, predator, and year effects. Journal of Wildlife Management 70:1257-1265.
Mahony, N., P.G. Krannitz and K. Martin (2006). Seasonal fecundity of sagebrush Brewer’s sparrows at the northern edge of the range. The Auk 123:512-523.
Hannon, S.J., and K. Martin (2006). Ecology of juvenile grouse during the transition to adulthood Journal of Zoology 289:422-433. (Invited)
Wiebe, K.L., W.D. Koenig, and K. Martin (2006). Evolution of clutch size in cavity-excavating birds: the nest site limitation hypothesis revisited American Naturalist 167:343-353.
Martin, K., A.R. Norris, and M.C. Drever (2006). Effects of bark beetle outbreaks on avian biodiversity in Interior British Columbia British Columbia Journal of Ecosystems Management 7:10-24.
Martin, K. (2005). Aspen: Prime real estate for nesting birds and mammals in Interior Douglas-fir forest in British Columbia Sustainable Forest Management Network advertisement in: Logging and Sawmilling Journal, May, page 51.
Martin, K (2005). Wildlife biodiversity and MPB [mountain pine beetle] in interior forests Branchlines 16:9-10
Robinson, P.A., A.R. Norris and K. Martin (2005). Interspecific nest sharing for Red-breasted Nuthatches and Mountain Chickadees in British Columbia, Canada Wilson Bulletin 117(4): 400-402.
Sandercock, B.K., K. Martin and S.J. Hannon (2005). Life history evolution in extreme environments: comparative demography of arctic and alpine ptarmigan Ecology 86: 2176-2186
Er, K.B.H, J.L. Innes, K. Martin, and B. Klinkenburg (2005). Forest loss with urbanization predicts bird extirpations in Vancouver Biological Conservation 126, 410-419.
Wilson, S. and K. Martin (2005). High elevation habitat associations of birds during fall migration Ecoscience 12:561-568.
Martin, K. and K. L. Wiebe (2004). Coping mechanisms of alpine and arctic breeding birds: extreme weather and limitations to reproductive resilience Integrative and Comparative Biology 44:177-185.
Leupin, E.E., T. E. Dickinson, and K. Martin (2004). Resistance of songbirds to habitat perforation in a high elevation conifer forest Canadian Journal of Forest Research 34:1919-1928
Lindh, H. and K. Martin (2004). Utility of systematic reserve selection approaches for conservation of North American mountain resort communities: a case study in Whistler, Canada Mountain Research and Development 24: 319-326
Aitken, K.E.H. and K. Martin (2004). Nest cavity availability and selection in aspen-conifer groves in a grassland landscape Canadian Journal of Forest Research 34:2099-2109
Martin, K., G. Brown, and J.R. Young (2004). The historic and current distribution of the Vancouver Island White-tailed Ptarmigan (Lagopus leucurus saxatilis) Journal of Field Ornithology 75:239-256
Melles, S., S. Glenn, and K. Martin (2003). Urban bird diversity and landscape complexity: Species–environment associations along a multiscale habitat gradient. Conservation Ecology 7(1): 5. [online]
Martin, K. (2003). Staying on top: Persistence and connectivity processes for birds in mountain habitats. The Banff Centre In: Martin, K. and D.S. Hik eds. The state of ecological and earth sciences in mountain areas. Science Highlights from symposium on ecological and earth sciences in mountain areas, September 2002, Banff, Alberta, Canada. Pp. 21-22.
Martin, K. and D.S. Hik (2003). The state of ecological and earth sciences in mountain areas. Ther Banff Centre Science Highlights from symposium on ecological and earth sciences in mountain areas, September 2002, Banff, Alberta, Canada. 46 pages.
Taylor, L., K. Martin, D.S. Hik and A. Ryall (editors) (2002). Ecological and Earth Sciences in Mountain Areas conference. Banff, AB. The Banff Centre 296 Pages.
Martin, K. (2002). Protocol for monitoring birds in high elevation habitats during fall migration Department of Forest Sciences, University of British Columbia and Candian Wildlife Service, Pacific and Western Region. 14 pp.
Aitken, K.E.H., K.L. Wiebe, and K. Martin (2002). Nest-site reuse patterns for a cavity-nesting bird community in interior British Columbia. The Auk 119: 391-402
Braun, C. E. and K. Martin (2001). Unusual summer plumage of white-tailed ptarmigan. Wilson Bulletin 113: 373-377
Martin, K. (2001). Wildlife communities in alpine and subalpine habitats. Oregon University Press, Corvallis. Chapter 9: pp. 239-260 in D.H. Johnson and T.A. O’Neil (Managing Directors), Wildlife-habitat Relationships in Oregon and Washington.
Martin, K., C. Doyle, S.J. Hannon, and F. Mueller. (2001). Forest grouse and ptarmigan. Oxford University Press. Chapter 11: pp. 240-260 in C.J. Krebs, S. Boutin and R. Boonstra (eds.), Ecosystem Dynamics of the Boreal Forest: The Kluane Project.
Martin, K. and L. Forbes (2001). Vancouver Island White-tailed Ptarmigan species account BC Ministry of Environment, Lands and Parks, Victoria, BC. 10p.
Congdon, B.C., J.F. Piatt, K. Martin, and V.L. Friesen. (2000). Mechanisms of population differentiation in marbled murrelets: Historical versus contemporary processes. Evolution 54:974-986.
Hochachka, W.M., K. Martin, F. Doyle, and C.J. Krebs. (2000). Censusing vertebrate populations using observational data. Canadian Journal of Zoology 78:521-529.
Martin, K., P.B. Stacey and C.E. Braun. (2000). Recruitment, dispersal, and demographic rescue in spatially-structured white-tailed ptarmigan populations. Condor 102:503-516.
Wiebe, K.L. and K. Martin. (2000). The use of incubation behaviour by ptarmigan to adjust reproductive investment costs after egg laying. Behavioural Ecology and Sociobiology 48:463-470.
Steidl, R. J. and L. Thomas (K. Martin, supervisor) (2000). Power analysis and experimental design. Oxford University Press. In press. In Scneiner, S. M. and J. Gurevitch (eds.) Design and Analysis of Ecological Experiments, 2nd edition.
Wiebe, K.L. and K. Martin (2000). The use of incubation behaviour by ptarmigan to adjust reproductive investment costs after egg laying Behavioural Ecology and Sociobiology, 48: 463-470
Martin, K. and J.M. Eadie. (1999). Nest Webs: a community wide approach to the management and conservation of cavity nesting birds. Forest Ecology and Management 115: 243-257. Abstract
Artiss, T., W.M. Hochachka, and K. Martin. (1999). Female foraging and male vigilance in white-tailed ptarmigan (Lagopus leucurus): opportunism or behavioural coordination? Behavioural Ecology and Sociobiology, 46: 429-434.
Boulanger, J., K. Martin, G. Kaiser and, A.E. Derocher. (1999). Evaluating uncertainty in estimating population trends for marbled murrelets. Society of Canadian Ornithologists, Special Pub. No. 1. Biology and Conservation of Forest Birds. pp. 53-63.
Dragon, S., C. Carey, K. Martin, and R. Baumann. (1999). Effect of high altitude and in vivo adenosine/b-adrenergic receptor blockade on ATP and 2,3BPG concentrations in red blood cells of avian embryos. Journal of Experimental Biology, 202:2787-2795. Abstract
Nelson, K.J. and K. Martin. (1999). Thermal aspects of nest-site location for vesper sparrows and horned larks in the Chilcotin grasslands, British Columbia. Studies in Avian Biology 19:137-143.
Vanderkist, B.A., X.H. Xue, R. Griffiths, K. Martin, W. Beauchamp, and T.D. Williams. (1999). Evidence of male-bias in capture samples of Marbled Murrelets from genetic studies in British Columbia. Condor 101:398-402.
Easton, W.E. and K. Martin. (1998). The effect of vegetation management on breeding bird communities in British Columbia. Ecological Applications 48:1092-1103.
Hannon, S.J., P.K. Eason and K. Martin. (1998). Willow Ptarmigan Philadelphia: The Academy of Natural Sciences; Washington, DC: The American Ornithologists’ Union, pp. 1-28. In: The Birds of North America, No. 369 (A. Poole and F. Gill, Eds.).
Krebs, C.J., A.R.E. Sinclair, R. Boonstra, S. Boutin, K. Martin, and J.N.M. Smith. (1998). Community dynamics of vertebrate herbivores – how can we untangle the web? in Herbivores: Between Plants and Predators. Editors: H. Olff, V.K. Brown, and R.H. Drent. pp.447-473, Chapter 15, Blackwells, Oxford.
Turkington, R.T., E. John, C.J. Krebs, M. Dale, V. Nams, R. Boonstra, S. Boutin, K. Martin, A.R.E. Sinclair and J.N.M. Smith. (1998). The effects of NPK fertilization for nine years on boreal forest vegetation in northwestern Canada. Journal of Vegetation Science 9:333-346.
Wiebe, K.L. and K. Martin. (1998). Age-specific patterns of reproduction in white-tailed and willow ptarmigan (Lagopus leucurus and L. lagopus). Ibis 140:14-24.
Wiebe, K.L. and K. Martin. (1998). Costs and benefits of nest cover for ptarmigan: Changes within and among years. Animal Behaviour 56:1137-1144.
Martin, K. and S. Ogle. (1998). The use of alpine habitats by fall migrating birds on Vancouver Island. Report, Department of Forest Sciences, University of British Columbia and Canadian Wildlife Service, Pacific and Western Region, Delta,B.C. 19 p. Posted online
Carey, C. and K. Martin. (1997). Physiological ecology of incubation of ptarmigan eggs at high and low altitudes. Wildlife Biology 3:211-218.
Holt, R.F. and K. Martin. (1997). Landscape modification and patch selection: the demography of two secondary cavity-nesting species colonizing clear-cuts. The Auk 114:443-455.
Wiebe, K.L. and K. Martin. (1997). Effects of predation, body condition and temperature on incubation rhythms of ptarmigan. Wildlife Biology 3: 218-227.
Hannon, S.J. and K. Martin. (1996). Mate fidelity and divorce in ptarmigan: polygyny avoidance on the tundra. Oxford University Press, London, U.K. pp. 192-210 in J.M. Black (ed.), Partnerships in Birds, the Ecology of Long-Term Monogamy
Thomas, L.J. and K. Martin. (1996). The importance of analysis method for the breeding bird survey population trend estimates. Conservation Biology 10:479-490.
C.E. Braun, K. Martin and L.A. Robb. (1993). White-tailed Ptarmigan Philadelphia: The Academy of Natural Sciences; Washington, DC: The American Ornithologists’ Union. In: The Birds of North America, No. 68 (A. Poole, and F. Gill, Eds.). 22 pages.